HLA-DR1, HLA-DR15, HLA-DR16 citrullination outside of cell?

HLA-DR1, HLA-DR15, HLA-DR16

HLA-DR1, gingivalis and RA
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3467347/

RA and citrullination
http://www.news-medical.net/news/20131031/Scientists-unravel-the-major-cause-of-protein-citrullination-in-rheumatoid-arthritis.aspx

P. Gingivalis and citrullination
http://www.roadback.org/forums/topic/pathogens-and-seropositive-ra/
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4131749/

smoking and RA both have citrullination
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4284707/

smoking HLA-DR? and citrullination
https://www.ncbi.nlm.nih.gov/labs/articles/22537824/

DR-15 and aspergillus
http://www.sciencedirect.com/science/article/pii/S1569199302000334

HLA- DR1, DR-15 and HPV
https://www.ncbi.nlm.nih.gov/pubmed/17412975

HPV has citrullinated proteins in the viral coat
https://www.ncbi.nlm.nih.gov/pubmed/18511237

HLA-dr15 and early age MS
https://www.ncbi.nlm.nih.gov/pubmed/10939572

eczema and MS
http://www.pathologyjournal.rcpa.edu.au/article/S0031-3025(16)35289-8/pdf
https://www.ncbi.nlm.nih.gov/pubmed/11903117

which seems to hint that early onset MS is not a mycobacteria but a different infection like staph?

Flaggrin, citrullinatin, and eczema
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3378480/